Information Package / Course Catalogue
Soil Enzymes
Course Code: ZTO618
Course Type: Area Elective
Couse Group: Third Cycle (Doctorate Degree)
Education Language: Turkish
Work Placement: N/A
Theory: 2
Prt.: 2
Credit: 3
Lab: 0
ECTS: 7
Objectives of the Course

The aim of this course is to teach to students defining reaction mechanisms of microbial enzymes and understanding relationships between soil enzymes and soil quality.

Course Content

Structure and classification of soil microbial enzymes, enzymes involved in phosphorus, carbon, sulphur and nitrogen-metabolism.

Name of Lecturer(s)
Learning Outcomes
1.Able to define structure, reactions, localizations and importance of soil enzymes
2.Able to connect between soil quality and soil microbial enzymes
3.Able to make a list of the applications affecting microbial enzyme activity
4.Capable of defining the role of soil enzymes in soil-plant relationships
5.Able to apply soil enzyme analyses and evaluate the results
Recommended or Required Reading
1.Soil Enzymes. Burns, R.G., 1978. Academic Press, London-New York-San Francisco. Shukla, G., Varma, A. 2011.
2.Selected soil enzymes; Examples of their potential roles in the ecosystem. Makoi, J.H.J.R., Ndakidemi, PA., 2008. Afrıcan J. Biotechn. 7: 181-191 .
3.Soil Microbial Ecology. Metting, F.B., Marcel Dekker, Inc. New York, Basel, Hong Kong, (1992).
4.Soil Enzymology. Springer. Berlin Heidelberg New York. Schinner, F., Öhlinger, R., Kandeler, E., Margesin, R., 1995.
5.Methods in Soil Biology. Springer. Berlin Heidelberg New York.
6.Soil Microbiology, Ecology and Biochemistry. 2006. by Eldor Alvin Paul, Elsevier Science, 552pp
7.Methods in Applied Soil Microbiology and Biochemistry. 1995. Kassem Alef, Paolo Nannipieri Academic Pres, 576 pp
Weekly Detailed Course Contents
Week 1 - Theoretical
Introduction and historical perspective
Week 1 - Preparation Work
Literature research
Week 2 - Theoretical
Structure and reaction mechanisms of soil enzymes
Week 2 - Preparation Work
Literature research
Week 3 - Theoretical
The factors affecting soil enzyme activity
Week 3 - Preparation Work
Literature research
Week 4 - Theoretical
Origin of enzymes in soil
Week 4 - Preparation Work
Literature research
Week 5 - Theoretical
Classification of enzymes
Week 5 - Preparation Work
Literature research
Week 6 - Theoretical
Enzymes involved in N-metabolism
Week 6 - Preparation Work
Literature research
Week 7 - Theoretical
Enzymes involved in C-metabolism
Week 7 - Preparation Work
Literature research
Week 8 - Intermediate Exam
Midterm Exam
Week 9 - Theoretical
Enzymes involved in P-metabolism
Week 9 - Preparation Work
Literature research
Week 10 - Theoretical
Enzymes involved in S-metabolism
Week 10 - Preparation Work
Literature research
Week 11 - Theoretical
Enzymes involved in intracellular metabolism
Week 11 - Preparation Work
Literature research
Week 12 - Theoretical
Role of enzymes in soil-plant relationships.
Week 12 - Preparation Work
Literature research
Week 13 - Theoretical
Role of enzymes in soil-plant relationships.
Week 13 - Preparation Work
Literature research
Week 14 - Theoretical
Enzymes as indicator of soil quality
Week 14 - Preparation Work
Literature research
Week 15 - Theoretical
Practice Examination
Week 16 - Final Exam
Final Exam
Assessment Methods and Criteria
Type of AssessmentCountPercent
Midterm Examination1%40
Final Examination1%60
Workload Calculation
ActivitiesCountPreparationTimeTotal Work Load (hours)
Lecture - Theory140228
Lecture - Practice140228
Seminar102525
Laboratory80216
Midterm Examination102525
Final Examination105050
TOTAL WORKLOAD (hours)172
Contribution of Learning Outcomes to Programme Outcomes
PÇ-1
PÇ-2
PÇ-3
PÇ-4
PÇ-5
OÇ-1
5
4
2
5
3
OÇ-2
5
5
4
5
4
OÇ-3
5
5
2
5
4
OÇ-4
5
4
5
4
5
OÇ-5
5
4
5
4
5
Adnan Menderes University - Information Package / Course Catalogue
2026